The purpose of the article is to determine and ground the organizational and pedagogical conditions of the Higher Military Educational Institutions special subjects lecturers professional competence development in post-graduate education system. The article contains the analysis of scientific researches concerning the determination of the next concepts: "pedagogical conditions", "organizational and pedagogical conditions". The conducted analysis of the scientific works indicates, that the concept "condition" is widespread in the modern scientific researches, with the view to practicality and optimization of educational process. The content of the concept "organizational and pedagogical conditions" is grounded. It presupposes sufficient and necessary circumstances and factors, allowing most beneficial environment of effective Higher Military Educational Institutions special subjects lecturers professional competence development pedagogical model functioning in post-graduate education system to achieve planned educational goals. Educational and pedagogical conditions, aimed at the unification of special subjects lecturers course training, professional competence development model functioning, taking into account the actual state of lecturers professional competence development, lecturers needs and requests during the process of educational content formation, which will satisfy the needs of pedagogical development as a subject of pedagogical practice.